Dedicated in memory of Stan Rand, 1932-2005.
Your frogs and friends in Panama miss you.
Ashley MacDonald, Marie Bottomley, Andrew Durso, Elizabeth Nixon, Thao Quan & John Pickering,
University of Georgia, Athens.
Taxonomy provided by
American Museum of Natural History, Amphibian Species of the World.
Compiled from Herps of Panama ,
Anurans of Las Cruces Biological Station, Costa Rica ,
Integrated Taxonomic Information Systems ,
The Amphibians and Reptiles of Costa Rica , Jay M. Savage, 2002,
Amphibian Species of the World , Edited by Darrel R. Frost, 1985,
and Amphibians and Reptiles of La Selva, Costa Rica, and the Caribbean Slope , Craig Guyer & Maureen A. Donnelly, 2005.
Where taxonomy differed between sources, the AMNH taxonomy was used.